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Utawarerumono is a story centering around the masked protagonist, Hakuoro, who one day is found by a family of two girls and their grandmother in a nearby forest close to their village. He is badly injured and is soon found to have amnesia, so they take him to their home and treat him until he is well again. Hakuoro is soon accepted into their village where he stays and lives with them, but before long Hakuoro finds that life in this village is being oppressed by the greedy emperor who rules over the country their village resides in. Soon after, these negative actions taken towards their village result in him leading a rebellion against the emperor of the country which later lands him to become emperor of a vast new country which is named Tusukuru, after the well-respected woman who helped save his life.
After becoming emperor and things begin to calm down, Hakuoro soon finds out that peace is hard to maintain and finds himself constantly in battle in order to protect the peace of his country and his people. Eventually, he is led into many bloody battles in order to fight for the freedom of all those residing in Tusukuru. Along the way, Hakuoro meets several other strong warriors from several other countries and tribes who are fully accepted into Hakuoro's growing family. Many moments of hardship and laughter ensue as time continues but with Hakuoro leading the way, all others are confident in his ability to lead.
The story's genre is at first a fantasy-style story with heavy Ainu influence, though later develops science fiction themes towards the end. Indeed, it first appears to take place in a fantasy world full of magical beings and interesting new species of humans, but it is later revealed that it takes place in Earth's distant future.

In the present age, an archaeological expedition unearths a mysterious fossil, a fossil of a disfigured beast. Unknown to the archaeologist, the fossil is the embodiment of a forgotten god, known as Witsuarunemitea. A superior scientist stumbles in, and after some confrontation, the mysterious stranger explains that humanity isn't ready for this Missing Link, the gap in evolution between men and apes. To prevent him from talking, he shoots the archaeologist. As the archaeologist slowly dies, his blood touches the fossil, and the forgotten god awakens.
Witsuarunemitea, a shattered and forgotten deity, sees opportunity in the dying archaeologist. He offers the archaeologist life, in exchange for becoming his vessel, so that Witsuarunemitea can finally find the sleep he so desires. Time passes, and eventually he becomes frozen in ice.
With Witsuarunemitea still bound in sleep as his mask, the archaeologist re-awakens many years into the future. He finds himself in an underground research facility, where he is being studied as an ancient human specimen. The archaeologist learns from the researchers that humanity is now unable to exist outside of a greenhouse environment, and hence the remnants of humanity now live underground.
Mizushima, a researcher, is assigned to the care and study of the archaeologist, who is now referred to as the Iceman. Upon discovering that the 'Iceman' can withstand the bacteria on Earth's surface, the researchers look to harness his unique genome and emulate the invigorating characteristics of his mask. Of the thousands of creations, only a select few prove to be viable, including experiments 3510 and 63. Experiment number 3510, a beautiful young girl, is given the name Mikoto by the Iceman; as is experiment number 63, a girl with black wings, whom is given the name Mutsumi. Mutsumi, acknowledging she has come from the Iceman's DNA, addresses him as "father".
The genome project encounters some problems, however, and the researchers in charge decide to re-freeze the Iceman for later study. Mizushima, aware of the human side of the Iceman, urges him to run, and gives him the facility's master key, a plain-colored ring recognized by the facility's doors.
The Iceman runs from the facility, taking Mikoto with him, whom he has fallen in love with. After a few years have passed, Mikoto gives birth to a daughter. The researchers, however, put on hazmat suits in order to pursue the Iceman on the surface world and recapture both him and Mikoto. Mikoto is dissected for further research, but while Iceman is being prepared to be frozen, his rage awakens Witsuarunemitea.
Outraged by the treatment of Mikoto, a new side of Witsuarunemitea begins to form. Filled with rage, Witsuarunemitea destroys most of the facility. Appalled by what the humans would do to obtain immortality, Witsuarunemitea grants their plea, transforming them into immortal gelatinous beings. Eventually, the great god accrues so much rage as to split himself into two halves: one which seeks destruction, and another, which seeks only to be destroyed.
Suddenly, Mutsumi appears, and empowered by Witsuarunemitea's divine blood, she puts both halves of Witsuarunemitea to sleep. Meanwhile, the few successful genetic experiments from the research facility begin to repopulate the Earth's surface, as the original humanity slowly dies out and is forgotten. The research facility is sealed away forever, and the people are told it is Witsuarunemitea's holy land.
Years later, a shift in the world awakens the two halves of Witsuarunemitea, who begin to combat each other; the destructive side wanting to help the populace of the world reach new height through wars, and its opposite through peaceful cooperation. The empress of Kunnekamun (whose people; the Shakukopuro, are being massacred) seeks out the destructive side, and is granted with a power called the Avu-Kamuu, massive bio-mechanical constructs covered in platemail, allowing them to exist in the world of war and hegemony.
Eventually, the fighting of the two halves causes a great earthquake. Arurū, a young girl of Kenashikourupe, is in the process of climbing a tree to recover a beehive when the earthquake strikes, and falls, becoming greatly injured. Erurū, Arurū's sister, prays Witsuarunemitea to heal Arurū, and promises, in exchange, her body and soul. Witsuarunemitea obliges.
Both halves of Witsuarunemitea are heavily injured in the battle, and both halves are put to sleep. The destructive half was found by Dii, a scholar of Onkamiyamukai, who fuses with the demigod. The half that wishes to be destroyed retains his body, and is found in the woods by Erurū.
The events of Utawarerumono then take place.


In Utawarerumono, there is a wide range of characters from several different countries that appear throughout the story. The main characters are all connected back to the male protagonist, Hakuoro, a man found injured in a forest who wears an irremovable mask and has lost his memories. He is a charismatic man who has the power to rally people under him for a cause when it comes time to fight. After earning much of the respect from the people who served under his command, Hakuoro was appointed emperor of the newly founded country Tusukuru, named after the grandmother of the woman who initially found him - Erurū. Erurū is a physician in training who often helps Hakuoro in much the same way a secretary or even a mother would. She has a forceful, but caring personality which is how she acts most of the time, especially around her younger sister Arurū who she cares for deeply. Arurū is a playful young girl who is shy and quiet, usually only saying a word or two when spoken to. After the god of the forest, Mutikapa is killed, Arurū raises her child as if it were her own, naming it Mukkuru. Since Erurū was the one who initially cared for him, Hakuoro regards her and her family the closest thing to a family of his own.
After Tusukuru is founded, Hakuoro gets many warriors under his command. The first is an impulsive and zealous dual-sword wielding former-bandit named Oboro. Dori and Gura are his two twin archer servants. Oboro has a younger sister named Yuzuha who has been sick with an unknown illness since birth, and is usually bedridden because of it. The next man to join Hakuoro is Benawi, a strong warrior who carries with him a calm and logical personality, though often gets irritated when Hakuoro ignores his duties as a ruler. His right hand man is Kurou, whose strength is comparable to Oboro's. There are several strong women who fight for Hakuoro as well. Urutori of the winged Onkamiyamukai race comes to Tusukuru as a priestess sent to look out for the country's welfare. She, and her younger sister Kamyu who follows Urutori to Tusukuru, have powerful magic abilities akin to those of their race. Karurauatsūrei, whose name is shortened to Karura for simplicity, is found having slaughtered many soldiers from another country who were holding her prisoner. Hakuoro decides to have her join his cause and she agrees. Karura carries a ridiculously heavy, black personalized blade only she can wield and has an affinity towards sake. Touka of the Evenkuruga tribe initially fought against Hakuoro under a false pretense though later joins him after her previous lord Orikakan is murdered by another emperor, Niwe, a common enemy of Touka and Hakuoro. Touka is very protective of Hakuoro, acting as his bodyguard. She usually has a serious personality though sometimes she is shown to have a tsundere personality.
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